1. Hawai'i Institute of Marine Biology University of Hawai'i Kāne'ohe HI 96744 USA
2. School of Life Sciences, College of Medical, Veterinary and Life Sciences University of Glasgow Glasgow G12 8QQ UK
3. Red Sea Research Center Division of Biological and Environmental Science and Engineering King Abdullah University of Science and Technology (KAUST) Thuwal 23955‐6900 Saudi Arabia
4. Department of Environment and Agriculture Curtin University PO Box U1987 Perth WA 6845 Australia
5. Section of Ichthyology California Academy of Sciences San Francisco CA 94118 USA