1. Department of Biosciences; College of Life and Environmental Sciences; University of Exeter; Exeter Devon U.K
2. Centre for Ecology and Hydrology Dorset; Winfrith Technology Centre; Dorchester Dorset U.K
3. The School of Biological and Chemical Sciences; Queen Mary University of London; London U.K
4. Game and Wildlife Conservation Trust (GWCT) - Salmon and Trout Research Centre; The River Laboratory; East Stoke Wareham Dorset U.K