1. Macroevolution & Macroecology, Division of Ecology & Evolution, Research School of Biology; Australian National University; Canberra ACT 2601 Australia
2. Department of Biochemistry and Molecular Biology, Bio21 Molecular Science and Biotechnology Institute; The University of Melbourne; Melbourne VIC 3010 Australia
3. School of Life and Environmental Sciences; University of Sydney; Sydney NSW 2006 Australia