1. Department of Plant Biology and Forest Genetics; Uppsala BioCenter; Swedish University of Agricultural Sciences and Linnean Center for Plant Biology; PO Box 7080; Uppsala; SE-75007; Sweden
2. Departamento de Biologia Molecular y Bioquimica; Facultad de Ciencias; Universidad de Malaga; Campus de Teatinos; Malaga; 290071; Spain
3. Department of Crop Production Ecology; Swedish University of Agricultural Sciences and Linnean Center for Plant Biology; PO Box 7043; Uppsala; SE-75007; Sweden