1. Key Laboratory of Plant Molecular Physiology, Institute of Botany Chinese Academy of Sciences Beijing 100093 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Jiangsu Co‐Innovation Center for Modern Production Technology of Grain Crops Yangzhou University Yangzhou 225009 China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China