1. Hunan Provincial Key Laboratory of Protein Engineering in Animal Vaccines, College of Veterinary Medicine Hunan Agricultural University Changsha Hunan Province China
2. Hunan Co‐Innovation Center of Animal Production Safety, College of Veterinary Medicine Hunan Agricultural University Changsha Hunan Province China
3. Department of Zoology University of Swabi Swabi 23340 Khyber Pakhtunkhwa Pakistan