1. Department of Entomology; University of California; Davis CA 95616 USA
2. Center for Population Biology; University of California; Davis CA 95616 USA
3. Hakubi Center for Advanced Research; Kyoto University; Kyoto 606-8501 Japan
4. Department of Natural Environmental Science; Niigata University; Niigata 950-2181 Japan
5. Department of Evolution and Ecology; University of California; Davis CA 95616 USA
6. Department of Environmental Science; University of Eastern Finland; Kuopio FIN-70211 Finland