1. Department of Animal and Plant Sciences; University of Sheffield; Sheffield; S10 2TN; UK
2. National Institute for Mathematical and Biological Synthesis; Claxton Building; 1122 Volunteer Boulevard; Suite 106; Knoxville; TN; 37996-3410; USA
3. Evolutionary Ecology Group; Goethe-University Frankfurt; Max-von-Laue-Strasse 13; 60438; Frankfurt am Main; Germany