1. School of Biological Sciences; University of East Anglia; Norwich Research Park, Norwich, Norfolk NR4 7TJ UK
2. Department of Zoology; Edward Grey Institute; University of Oxford; Oxford OX13PS UK
3. School of Biology; The Faculty of Biological Sciences, University of Leeds; Leeds LS2 9JT UK
4. Behavioural Ecology and Physiological Group; Groningen Institute for Evolutionary Life Sciences; University of Groningen; PO Box 11103, 9700 CC, Groningen The Netherlands
5. Department of Animal and Plant Sciences; University of Sheffield; Sheffield S10 2TN UK
6. Nature Seychelles; PO BOX 1310, Mahe Republic of Seychelles