1. Department of Watershed Sciences and the Ecology Center Utah State University Logan Utah USA
2. Native Fish Ecology and Conservation Program, Division of Science and Resource Management Grand Canyon National Park, National Park Service Flagstaff Arizona USA
3. U.S. Geological Survey, Utah Cooperative Fish and Wildlife Research Unit, Department of Watershed Sciences Utah State University Logan Utah USA
4. U.S. Geological Survey, Southwest Biological Science Center Grand Canyon Monitoring and Research Center Flagstaff Arizona USA
5. School of Environmental Science Simon Fraser University Vancouver British Columbia Canada
6. Upper Colorado Regional Office U.S. Bureau of Reclamation Salt Lake City Utah USA