1. Laboratory for Evolutionary Morphology RIKEN Center for Biosystems Dynamics Research (BDR) Kobe Hyogo Japan
2. Department of Biology, Graduate School of Science Kobe University Kobe Hyogo Japan
3. Aix‐Marseille Université, CNRS UMR 7288, IBDM Marseille France
4. Laboratory for Evolutionary Morphology RIKEN Cluster for Pioneering Research (CPR) Kobe Hyogo Japan