1. Key Laboratory of Algal Biology Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 P. R. China
2. University of Chinese Academy of Sciences Beijing 100039 P. R. China
3. CAS Key Laboratory of Marine Ecology and Environmental Sciences Institute of Oceanology Chinese Academy of Sciences Qingdao 266071 P. R. China
4. School of Life Science Shanxi University Taiyuan 030006 P. R. China
5. State Key Laboratory of Freshwater Ecology and Biotechnology Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 P. R. China