1. Centre for Ecology, Evolution & Biogeochemistry; EAWAG Federal Institute of Aquatic Science and Technology; Seestrasse 79 6074 Kastanienbaum Switzerland
2. Institute for Ecology and Evolution; University of Bern; Baltzerstrasse 6 3012 Bern Switzerland
3. Department of Biological Science; California State University Fullerton; Fullerton CA 92831 USA