1. College of Veterinary Medicine; Huazhong Agricultural University; Wuhan 430070 China
2. Tianjin Helaiente biological science and Technology Co., Ltd; Tianjin 301709 China
3. Key Laboratory of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education; College of Animal Science and Technology; Huazhong Agricultural University; Wuhan 430070 China
4. State Key Laboratory of Agricultural Microbiology; College of Veterinary Medicine; Huazhong Agricultural University; Wuhan 430070 China