1. CAS Key Laboratory of Tropical Marine Bio-resources and Ecology; Guangdong Key Laboratory of Marine Materia Medica; RNAM Center for Marine Microbiology; South China Sea Institute of Oceanology; Chinese Academy of Sciences; Guangzhou 510301 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Department of Chemical Engineering; Pennsylvania State University, University Park; PA 16802-4400 USA
4. Department of Biochemistry and Molecular Biology; Pennsylvania State University; University Park PA 16802-4400 USA