1. CAS Key Laboratory of Biofuels Shandong Provincial Key Laboratory of Synthetic Biology Qingdao Institute of Bioenergy and Bioprocess Technology Chinese Academy of Sciences Qingdao 266101 China
2. Dalian National Laboratory for Clean Energy Qingdao 266101 China
3. University of Chinese Academy of Sciences Chinese Academy of Sciences Beijing 100049 China
4. Department of Biotechnology and Enzyme Catalysis Institute of Biochemistry Greifswald University Felix-Hausdorff-Str. 4 D-17487 Greifswald Germany