1. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China
2. State Key Laboratory of Vegetation and Environmental Change (LVEC), Institute of Botany Chinese Academy of Sciences Beijing 100093 China
3. Department of Botany, National Museum of Natural History Smithsonian Institution PO Box 37012 Washington DC 20013‐7012 USA
4. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
5. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
6. Department of Biology—Ecoinformatics and Biodiversity Aarhus University 8000 Aarhus C Denmark
7. Key Laboratory of Horticultural Plant Biology (Ministry of Education), College of Horticulture and Forestry Science Huazhong Agricultural University Wuhan 430070 China
8. College of Architecture and Urban Planning Tongji University Shanghai 200092 China