1. Department of Ecology, Evolution, and Behavior University of Minnesota St Paul MN 55108 USA
2. Department of Biological Sciences Minnesota State University Mankato Mankato MN 56001 USA
3. Instituto de Investigaciones en Ecosistemas y Sustentabilidad Universidad Nacional Autónoma de México Antigua Carretera a Pátzcuaro No. 8701, Col. Ex Hacienda de San José de la Huerta Morelia Michoacán 58190 Mexico
4. Center for Tree Science and Herbarium The Morton Arboretum Lisle IL 60532 USA
5. The Field Museum 1400 S Lake Shore Drive Chicago IL 60605 USA