1. Department of Ecology and Evolutionary Biology; University of California; Santa Cruz California 95064 USA
2. School of Biological Sciences; University of Nebraska-Lincoln; Lincoln Nebraska 68588 USA
3. Station d'Ecologie Expérimentale du CNRS (USR2936); 2 route du CNRS 09200 Moulis France
4. Institute for Advanced Studies in Toulouse; Toulouse School of Economics; 21 allée de Brienne 31015 Toulouse France