1. Department of Biology; University of Eastern Finland; P.O. Box 111 Joensuu FI-80101 Finland
2. Institute for Systematic Botany; University of Zurich; Zollikerstrasse 107 Zurich CH-8008 Switzerland
3. Finnish Environment Institute; Friendship Park Research Centre; Lentiirantie 342 B Kuhmo FI-88900 Finland
4. National Museums of Scotland; Chambers Street Edinburgh EH1 1JF UK
5. Department of Natural Sciences; Tromsø University Museum; Tromsø 9037 Norway
6. Liinalammintie 11 as. 6; Turenki FI-14200 Finland