1. Department of Ecology and Evolutionary Biology University of Toronto Mississauga ON L5L 1C6 Canada
2. Forest Biodiversity Research Network, Department of Forest Ecosystems and Society Oregon State University Corvallis OR 97331‐5704 USA
3. Biodiversity Section Department of Natural Resources and Energy Development Fredericton NB E3C 2G6 Canada
4. Swiss Ornithological Institute Sempach CH‐6204 Switzerland
5. Department of Botany and Plant Pathology Oregon State University Corvallis OR 97331 USA
6. Smithsonian Tropical Research Institute Panama City 0843‐03092 Panama