1. Department of Genetic and Behavioral Neuroscience; Gunma University Graduate School of Medicine; 3-39-22 Showa-machi Maebashi Gunma 371-8511 Japan
2. Institute of Experimental Animal Research; Gunma University Graduate School of Medicine; Maebashi Japan
3. Japan Science and Technology Agency; Core Research for Evolutional Science and Technology (CREST); Sanbancho Chiyoda-ku Tokyo Japan