1. MRC/Wits Developmental Pathways for Health Research Unit; Department of Paediatrics; School of Clinical Medicine; University of the Witwatersrand; Johannesburg South Africa
2. Division of Human Genetics; School of Pathology; University of the Witwatersrand and National Health Laboratory Service; Johannesburg South Africa
3. Department of Paediatrics; University of Cambridge; Cambridge UK
4. Wellcome Trust-MRC Institute of Metabolic Science; Cambridge UK