1. Department of Biointeractions and Crop Protection Rothamsted Research Harpenden Hertfordshire AL5 2JQ UK
2. School of Animal, Rural and Environmental Sciences Nottingham Trent University Southwell NG25 0QF UK
3. Institute of Evolutionary Biology School of Biological Sciences The University of Edinburgh Edinburgh EH9 3JT UK
4. Department of Animal and Plant Sciences University of Sheffield Sheffield South Yorkshire S10 2TN UK