1. Laboratory of Protozoology; Guangzhou Key Laboratory of Subtropical Biodiversity and Biomonitoring; School of Life Science; South China Normal University; Guangzhou 510631 China
2. Microbial Ecology and Matter Cycle Group; School of Marine Sciences; Sun Yat-sen University; Zhuhai 519082 China
3. School of Ocean; Yantai University; Yantai 264005 China
4. Department of Life Sciences; Natural History Museum; Cromwell Road London SW7 5BD UK