1. Department of Plant Science; Plant Genomics and Breeding Institute; Research Institute of Agriculture and Life Sciences; Seoul National University; Seoul 151-921 South Korea
2. Department of Plant Life and Environmental Science; Hankyong National University; Ansung 456-749 South Korea
3. Department of Plant Molecular Systems Biotechnology; Crop Biotech Institute; Kyung Hee University; Yongin 446-701 South Korea
4. Crop Biotechnology Institute; GreenBio Science and Technology; Seoul National University; Pyeongchang 232-916 South Korea