1. Coastal and Marine Research Centre, Australian Rivers Institute Griffith University Nathan, QLD 4111 Australia
2. Centre for Tropical Water and Aquatic Ecosystem Research (TropWATER), College of Science and Engineering, Division of Tropical Environment and Societies James Cook University Townsville, QLD 4811 Australia
3. North Queensland Dry Tropics Natural Resources Management Townsville, QLD 4811 Australia