1. Department of Cellular and Molecular Medicine, Faculty of Health and Medical Sciences; University of Copenhagen; Copenhagen North Denmark
2. Department of Gastroenterology, Medical Section, Herlev Hospital; University of Copenhagen; Copenhagen Denmark
3. Department of Molecular Disease Biology, Faculty of Health and Medical Sciences; University of Copenhagen; Copenhagen East Denmark
4. Department of Biology, Faculty of Science; University of Copenhagen; Copenhagen Denmark
5. School of Biological Sciences; University of East Anglia; Norwich UK
6. Section of Oncopathology and Regenerative Biology, Department of Pathology, Faculty of Medicine; University of Miyazaki; Miyazaki Japan
7. Proteases and Tissue Remodeling Section, National Institute of Dental and Craniofacial Research; National Institutes of Health; Bethesda Maryland