1. U.S. Geological Survey; National Research Program; Boulder CO 80303 USA
2. Center for Limnology; University of Wisconsin-Madison; 680 N. Park St. Madison WI 53706 USA
3. St. Olaf College; Northfield MN 55057 USA
4. Department of Ecology, Evolution and Behavior; University of Minnesota; St. Paul MN 55108 USA