1. Department of Marine Science Incheon National University Incheon South Korea
2. Research Institute of Basic Sciences Incheon National University Incheon South Korea
3. Jiangsu Key Laboratory of Marine Bioresources and Environment Jiangsu Ocean University Lianyungang China
4. Department of Ecology and Evolutionary Biology University of Connecticut Stamford Connecticut USA
5. Fisheries Seed and Breeding Research Institute National Institute of Fisheries Science Jeonnam Korea