1. Tianjin Key Laboratory of Animal and Plant Resistance/College of Life Science; Tianjin Normal University; Tianjin 300387 China
2. Tianjin Center for Control and Prevention of Aquatic Animal Infectious Disease; Tianjin 300221 China
3. Louisiana State University Agricultural Center; Louisiana State University; Baton Rouge Louisiana 70820 USA
4. School of Renewable Natural Resources; Louisiana State University; Baton Rouge Louisiana 70820 USA