1. Department of Biology Utah State University Logan UT 84322 USA
2. EBIO University of Colorado Boulder Boulder CO 80309 USA
3. Department of Biological Sciences University of Notre Dame Notre Dame IN 46556 USA
4. CEFE Univ Montpellier Montpellier France
5. CNRS, EPHE, IRD Univ Paul Valéry Montpellier 3 Montpellier France