1. Centre for Ecology and Conservation; University of Exeter; Penryn Campus Cornwall TR10 9FE UK
2. Environment and Sustainability Institute; University of Exeter; Penryn Campus Cornwall TR10 9FE UK
3. Department of Zoology; School of Natural Sciences; Trinity College Dublin; Dublin 2 Ireland
4. Department of Biosciences; University of Helsinki; PO Box 65 (Viikinkaari 1) Helsinki FI-00014 Finland
5. Centre for Research in Ecology, Evolution and Behaviour; University of Roehampton; London SW15 4JD UK