1. Department of Biomedical Science and Environmental Biology, College of Life Science Kaohsiung Medical University Kaohsiung Taiwan
2. Mindanao State University‐Malabang Community High School Malabang Lanao del Sur Philippines
3. Graduate Institute of Medicine, Kaohsiung Medical University Kaohsiung Taiwan
4. Department of Biological Sciences and Environmental Studies, College of Science and Mathematics University of the Philippines Mindanao Davao City Philippines
5. Systematic Zoology Laboratory, Department of Biological Sciences Tokyo Metropolitan University Tokyo Japan
6. Department of Ecology and Evolutionary Biology, Biodiversity Institute University of Kansas Lawrence Kansas USA