1. Biodiversity Research Center Academia Sinica Taipei Taiwan
2. Mississippi Entomological Museum, Department of Entomology and Plant Pathology Mississippi State University Mississippi State MS U.S.A.
3. Department of Integrative Ecology Estación Biológica de Doñana (EBD‐CSIC) Seville Spain
4. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor MI U.S.A.