1. Institute of Water Sciences and Technology, Postgraduate Programs in: Society, Environment and Quality of Life; Biodiversity; Society, Nature and Development Federal University of Western Pará, Unidade Tapajós Santarém Pará 68040‐255 Brazil
2. Centre of Marine Sciences University of Algarve, Campus de Gambelas Faro 8005‐139 Portugal
3. Postgraduate Program in Ecology, Ecology and Conservation Laboratory, Institute of Biological Sciences Federal University of Pará Belém Pará 66075‐110 Brazil