1. Department of Biological Sciences; Graduate School of Science; University of Tokyo; 7-3-1 Hongo Bunkyo-ku Tokyo 113-0033 Japan
2. RIKEN Center for Sustainable Resource Science; 1-7-22 Suehiro-cho, Tsurumi-ku Yokohama-shi Kanagawa 230-0045 Japan
3. Department of Natural Science; Faculty of Science; Kochi University; 2-5-1 Akebono-cho Kochi-shi 780-8520 Japan
4. Graduate School of Integrated Arts and Sciences; Doctoral Course; Kuroshio Science; Kochi University; 2-5-1 Akebono-cho Kochi-shi 780-8520 Japan