1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences Guangzhou China
2. Key Laboratory of Genetics and Germplasm Innovation of Tropical Special Forest Trees and Ornamental Plants (Ministry of Education), College of Forestry, Hainan Univ. Haikou China
3. Biology Dept, Boston Univ. Boston MA USA
4. National Park Service, Acadia National Park and Schoodic Education and Research Center Bar Harbor ME USA