1. Department of Ecology and Evolutionary Biology; Cornell University; Ithaca NY 14853 USA
2. Division of Biological Sciences; University of Montana; Missoula MT 59812 USA
3. CMPG Lab; Institute of Ecology and Evolution; University of Bern; Baltzerstrasse 6 3012 Bern Switzerland
4. School of Natural Science; Hampshire College; Amherst MA 01002 USA