1. Department of Ecology and Evolutionary Biology; Brown University; Providence RI 02912 USA
2. The Josephine Bay Paul Center for Comparative Molecular Biology and Evolution; Marine Biological Laboratory; 7 MBL Street Woods Hole MA 02543 USA
3. Sea Education Association; P.O. Box 6 Woods Hole MA 02543 USA
4. Department of Earth Environmental and Planetary Sciences; Brown University; Providence RI 02912 USA