1. Oshima Branch Fisheries Laboratories Kinki University Oshima Kushimoto Wakayama Japan
2. Aquatic Resources Authority of Panama Panama Republic of Panama
3. Graduate School of Agriculture Kinki University Nakamachi Nara Japan
4. Inter‐American Tropical Tuna Commission La Jolla Shores Drive La Jolla CA USA
5. Inter‐American Tropical Tuna Commission Achotines Laboratory Las Tablas Los Santos Province Republic of Panama