1. National Education Minster Key Laboratory of Plant Genetic Improvement and Comprehensive Utilization; Fujian Agriculture and Forestry University; Fuzhou Fujian 350002 China
2. College of Life Science; Fujian Agriculture and Forestry University; Fuzhou Fujian 350002 China
3. Department of Life Science; Luoyang Normal University; Luoyang Henan 471022 China
4. Center for Plant Cell Biology, Institute for Integrative Genome Biology, Department of Botany and Plant Sciences; University of California at Riverside; Riverside CA 92521 USA