1. Naturalis Biodiversity Center P.O. Box 9517 2300 RA Leiden The Netherlands
2. Institute of Biology Leiden Leiden University P.O. Box 9505 2300 RA Leiden The Netherlands
3. Herbarium Bogoriense, Botany Division, Research Center for Biology Indonesian Institute of Sciences Cibinong Bogor Indonesia
4. North Gorua Village RT 006/RW 03, North Tobelo District, North Halmahera North Maluku Indonesia
5. Hortus botanicus Leiden University 2311 GJ Leiden The Netherlands
6. Bumi Mekar Wangi Residence Blok C6 No. 12, RT 04/RW 05, Tanah Sereal Bogor 16168 Indonesia
7. Missouri Botanical Garden St. Louis MO 63110 USA
8. Institut de Systématique, Évolution, et Biodiversité (ISYEB), Muséum National d'Histoire Naturelle, Centre National de la Recherche Scientifique Sorbonne Université, École Pratique des Hautes Études, Université des Antilles C.P. 39, 57 rue Cuvier Paris 75005 France
9. Department of Botany, MRC‐166, National Museum of Natural History Smithsonian Institution Washington DC 20013‐7012 USA