1. The Federal Research Center Institute of Cytology and Genetics SB RAS Novosibirsk 630090 Russia
2. L.K. Ernst Federal Research Center for Animal Husbandry Podolsk 142132 Russia
3. Novosibirsk State University Novosibirsk 630090 Russia
4. Russian State Agrarian University Moscow Timiryazev Agricultural Academy Moscow 127550 Russia
5. Royal Veterinary College University of London London NW1 0TU UK