1. School of Medical Sciences; Institute of Medical Sciences; University of Aberdeen; Aberdeen; AB25 2ZD; UK
2. CNRS-UMR 5175 Centre d'Ecologie Fonctionnelle et Evolutive; 1919 route de Mende; 34293; Montpellier Cedex 5; France
3. University Museums; King's College; University of Aberdeen; Aberdeen; AB24 3SW; UK
4. Institute of Genetics; University of Nottingham; Queen's Medical Centre; Nottingham; NG7 2UH; UK
5. Marine Biology & Ecology Research Centre; University of Plymouth; Plymouth; PL4 8AA; UK