1. Department of Biological Sciences and Marine Research Institute; University of Cape Town; Private Bag X3 Rondebosch 7701 South Africa
2. School of Plant Biology and Ocean Institute; University of Western Australia; Crawley WA 6009 Australia
3. Department of Biological Sciences; University of Cape Town; Rondebosch 7701 South Africa
4. Department of Biodiversity; University of Limpopo; Private Bag X1106 Sovenga 0727 South Africa
5. Fisheries Research; Department of Agriculture, Forestry and Fisheries; Private Bag X2 Roggebaai 8012 South Africa