1. College of Animal Science and Technology, Northwest A&F University; Shanxi Key Laboratory of Molecular Biology for Agriculture; Yangling 712100 China
2. Pearl River Fisheries Research Institute, Chinese Academy of Fishery Sciences; Key Laboratory of Fishery Drug Development, Ministry of Agriculture, Key Laboratory of Aquatic Animal Immune Technology, Guangdong Province; Guangzhou 510380 China
3. Freshwater Aquaculture Collaborative Innovation Center of Hubei Province; Wuhan 430070 China