1. School of Animal Biology; The University of Western Australia; 35 Stirling Highway Crawley WA 6000 Australia
2. Science and Conservation Division; Department of Parks and Wildlife; Bentley Delivery Centre; Locked Bag 104 Perth WA 6983 Australia
3. School of Plant Biology; The University of Western Australia; 35 Stirling Highway Crawley WA 6000 Australia
4. Centre for Evolutionary Biology; Centre of Excellence in Natural Resource Management; University of Western Australia; PO Box 5771 Albany WA 6332 Australia