1. Department of Botany and Plant Pathology Purdue University West Lafayette Indiana 47907 USA
2. Purdue Center for Plant Biology Purdue University West Lafayette Indiana 47907 USA
3. State Key Laboratory of Crop Genetics and Germplasm Enhancement, Center of Pear Engineering Technology Research Nanjing Agricultural University Nanjing Jiangsu 210095 China
4. Division of Biology and Biological Engineering California Institute of Technology Pasadena California 91125 USA
5. Howard Hughes Medical Institute California Institute of Technology Pasadena California 91125 USA