1. Department of Biology, Faculty of Science; University of Guilan; Rasht Iran
2. Fisheries Department, Faculty of Natural Resources; University of Guilan; Sowmeh-sara Iran
3. Fisheries Department, Faculty of Natural Resources and Marine Sciences; Tarbiat Modares University; Noor Iran
4. Instituto de Ciencias del Mar (CSIC); Passeig Marítim 37-49 E-08003 Barcelona Cataluña Spain